Dsungaripteroid pterodactyloid pterosaur Kimmeridgian age of the upper Jurassic the formation of Argiles d'Octeville at the Cap de la Heve near Ecqueville shellfish eater, volant piscivore found in a marine claystone layer fibrous texture like horn sheath the back of its head forever lost concave leading margin crested wave a subclade most inclusive of the lophocratia quite similar to the germanodactylidae abrupt beginning to a larger crest